Jane Goodall, the primatologist whose groundbreaking study of wild chimpanzees transformed science and inspired generations to see the natural world as kin, was remembered on Wednesday as both a visionary researcher and a moral force for the planet she devoted her life to protecting.

“She reminded us that change begins with compassion and that our humanity is our greatest tool,” said the actor and environmental activist Leonardo DiCaprio, one of several speakers who paid tribute to Dr. Goodall at her funeral at Washington National Cathedral. “She never lingered in despair. She focused on what could be done.”

An urn containing her ashes, draped in white, was placed inside the soaring Gothic cathedral as bells tolled solemnly over Northwest Washington. A stuffed chimpanzee clutching a ba

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