Gramma the Galápagos tortoise, oldest resident of San Diego Zoo, dies at about 141

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died Nov. 20.

It's not clear exactly when the tortoise arrived at the San Diego Zoo, but zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galápagos tortoises.

As the world changed around her, she delighted visitors with her sweet, shy personality. She lived through two World Wars and 20 U.S. presidents.

Her care specialists affectionately called her “the Queen of the Zoo.” She was suffering from bone conditions related to her old age