LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died Nov. 20.

It's not clear exactly when the tortoise arrived at the San Diego Zoo, but zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galapagos tortoises.

This photo provided by the San Diego Zoo Wildlife Alliance shows Gramma, a Galapagos tortoise and the oldest animal at the San Diego Zoo, eating a banana stalk May 17, 2023. San Diego Zoo Wildlife Alliance via AP

As the world changed around her, she delighted visitors with

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