Jeffery Holden takes notes over a wolf-killed moose on Isle Royale. (Anna Burke photo)

The cover of “Dead Moose on Isle Royale: Off Trail with the Citizen Scientists of the Wolf-Moose Project.” (Michigan State University Press)

Jeffery Holden takes notes over a wolf-killed moose on Isle Royale. (Anna Burke photo)

LANSING — Jeffrey Holden spends his summers knee-deep in dead moose.

In his new book “Dead Moose on Isle Royale: Off Trail with the Citizen Scientists of the Wolf-Moose Project” (Michigan State University Press, $24.95), Holden turns decades of volunteer field notes and short essays into an off-trail narrative about the people who sustain one of ecology’s longest-running studies.

The Wolf-Moose Project at Isle Royale National Park started with scientists from Purdue Universit

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